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Raptor 478 is piloted by Lt. Sharon "Boomer" Valerii with ECO Alex "Crashdown" Quartararo on at least two occasions. It flew during Lee "Apollo" Adama and Kara "Starbuck" Thrace's CAP to intercept Olympic Carrier (TRS: "33") and surveyed damage to Galactica's water storage tanks (TRS: "Water").
When CPO Galen Tyrol refers to a Raptor which Valerii had piloted on 47 missions, he may be referring to this one (TRS: "Flight of the Phoenix"). Valerii is also known to have piloted Raptor 312 (TRS: Miniseries).